Vibrissa below or at the level of lower margin of eye; head in profile subquadrate; vertical diameter of eye less than 1.5x the horizontal diameter; postocellar setae cruciate or so strongly converging that the tips nearly meet. Frons in male as wide as in female. Basisternum small, without precoxal bridge. Wing with subcostal notch seldom evident, never well-developed; R4+5 and M1 not or only slightly converging at tips; distal margin of anal cell (CuA2) usually rounded (if not, then epistoma extended dorsally, subtriangular). First tarsomere of hind leg often with a posteroventral comb of setae. Abdominal tergites of male without silvery microtomentum.